%PDF-1.6
%
1 0 obj
<>
endobj
2 0 obj
<>stream
Matthew T. Balazik, Daniel J. Farrae, Tanya L. Darden, Greg C. Garman
Genetic differentiation of spring-spawning and fall-spawning male Atlantic sturgeon in the James River, Virginia
www.plosone.org
www.plosone.org
endstream
endobj
3 0 obj
<>/XObject<>>>/CropBox[0 0 612 792]/MediaBox[0 0 612 792]/Parent 13 0 R/Annots 14 0 R/Contents 15 0 R/TrimBox[0 0 612 792]>>
endobj
14 0 obj
[16 0 R 17 0 R 18 0 R 19 0 R 20 0 R 21 0 R 22 0 R 23 0 R 24 0 R 25 0 R 26 0 R 27 0 R 28 0 R 29 0 R 30 0 R 31 0 R 32 0 R 33 0 R 34 0 R 35 0 R 36 0 R 37 0 R 38 0 R 39 0 R 40 0 R 41 0 R 42 0 R 43 0 R 44 0 R 45 0 R 46 0 R 47 0 R 48 0 R 49 0 R]
endobj
16 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref001)>>
endobj
17 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref002)>>
endobj
18 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref003)>>
endobj
19 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref004)>>
endobj
20 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref005)>>
endobj
21 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref001)>>
endobj
22 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref005)>>
endobj
23 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref006)>>
endobj
24 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref007)>>
endobj
25 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref008)>>
endobj
26 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref009)>>
endobj
27 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref010)>>
endobj
28 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref001)>>
endobj
29 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref005)>>
endobj
30 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref006)>>
endobj
31 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref010)>>
endobj
32 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref011)>>
endobj
33 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref004)>>
endobj
34 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref012)>>
endobj
35 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref013)>>
endobj
36 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref014)>>
endobj
37 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref015)>>
endobj
38 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref001)>>
endobj
39 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref004)>>
endobj
40 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref010)>>
endobj
41 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref016)>>
endobj
42 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref017)>>
endobj
43 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref018)>>
endobj
44 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref019)>>
endobj
45 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref010)>>
endobj
46 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref018)>>
endobj
47 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref020)>>
endobj
48 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref021)>>
endobj
49 0 obj
<>/Border[0 0 0]/A 50 0 R>>
endobj
50 0 obj
<>
endobj
15 0 obj
[51 0 R 52 0 R 53 0 R 54 0 R 55 0 R 56 0 R 57 0 R 58 0 R 59 0 R 60 0 R 61 0 R 62 0 R 63 0 R 64 0 R 65 0 R 66 0 R 67 0 R 68 0 R 69 0 R 70 0 R 71 0 R 72 0 R 73 0 R 74 0 R 75 0 R 76 0 R]
endobj
51 0 obj
<>stream
q
0.83 0.64 0.02 0 k
565.7953 674.9858 m
570.5575 674.9858 l
h
f*
200.0126 662.0031 m
204.7748 662.0031 l
h
f*
209.1969 662.0031 m
213.9591 662.0031 l
h
f*
218.4378 662.0031 m
223.2 662.0031 l
h
f*
227.622 662.0031 m
232.3843 662.0031 l
h
f*
548.2772 649.0205 m
553.0394 649.0205 l
h
f*
557.4614 649.0205 m
562.2236 649.0205 l
h
f*
566.7024 649.0205 m
571.4646 649.0205 l
h
f*
200.0126 635.9811 m
204.7748 635.9811 l
h
f*
209.1969 635.9811 m
213.9591 635.9811 l
h
f*
218.3811 635.9811 m
223.0866 635.9811 l
h
f*
227.5087 635.9811 m
237.0331 635.9811 l
h
f*
346.8472 622.9984 m
351.6094 622.9984 l
h
f*
356.0315 622.9984 m
360.7937 622.9984 l
h
f*
365.2724 622.9984 m
369.978 622.9984 l
h
f*
374.4567 622.9984 m
383.9811 622.9984 l
h
f*
448.1575 596.9764 m
457.6252 596.9764 l
h
f*
469.1906 571.011 m
473.8961 571.011 l
h
f*
478.3748 571.011 m
487.8992 571.011 l
h
f*
492.3213 571.011 m
501.8457 571.011 l
h
f*
442.4882 544.989 m
452.0126 544.989 l
h
f*
456.4346 544.989 m
465.9591 544.989 l
h
f*
449.4047 519.0236 m
454.1669 519.0236 l
h
f*
458.6457 519.0236 m
463.4079 519.0236 l
h
f*
283.2378 480.0189 m
292.7622 480.0189 l
h
f*
297.2409 480.0189 m
306.7087 480.0189 l
h
f*
311.1874 480.0189 m
320.7118 480.0189 l
h
f*
325.1339 480.0189 m
334.6583 480.0189 l
h
f*
427.6346 441.0142 m
437.1024 441.0142 l
h
f*
303.5339 414.9921 m
313.0583 414.9921 l
h
f*
317.4803 414.9921 m
327.0047 414.9921 l
h
f*
312.7181 172.9701 m
322.2425 172.9701 l
h
f*
556.8945 159.9874 m
566.4189 159.9874 l
h
f*
0 g
1 j
1 J
0 w
11.9999 0 0 11.9999 200.0125 706.0535 cm
BT
/F2 1 Tf
1 TL
-0.0033 Tc
0 0 Td
(Introduction)Tj
ET
Q
q
1 j
1 J
0 w
10 0 0 10 200.0125 689.1023 cm
BT
/F6 1 Tf
1 TL
-0.004 Tc
0 0 Td
(The)Tj
1.7801 0 Td
(Atlantic)Tj
3.4186 0 Td
(sturgeon)Tj
3.7247 0 Td
(\()Tj
/F11 1 Tf
[()30()1()10()4()2()-3()2()]TJ
4.3313 0 Td
[( )19(\n)-13()-9()12()6()13()18(\r)-4()]TJ
4.473 0 Td
[( )19(\n)-13()-9()12()6()13()18(\r)-4()]TJ
/F6 1 Tf
(,)Tj
4.7055 0 Td
(Acipenseridae\))Tj
6.2305 0 Td
(is)Tj
0.8447 0 Td
(an)Tj
1.2076 0 Td
(anadromous)Tj
5.2554 0 Td
(fish)Tj
-35.9714 -1.3039 Td
(that)Tj
1.7744 0 Td
(currently)Tj
3.8891 0 Td
(inhabits)Tj
3.4299 0 Td
(the)Tj
1.4684 0 Td
(east)Tj
1.7404 0 Td
(coast)Tj
2.2394 0 Td
(of)Tj
1.0148 0 Td
(North)Tj
2.6645 0 Td
(America)Tj
3.6397 0 Td
(from)Tj
2.2053 0 Td
(Labrador,)Tj
4.1386 0 Td
(CA)Tj
1.5647 0 Td
(to)Tj
1.0261 0 Td
(Florida,)Tj
3.3279 0 Td
(USA)Tj
2.1146 0 Td
([)Tj
0.83 0.64 0.02 0 k
(1)Tj
0 g
(,)Tj
0.83 0.64 0.02 0 k
-36.2378 -1.2982 Td
(2)Tj
0 g
(,)Tj
0.83 0.64 0.02 0 k
0.9184 0 Td
(3)Tj
0 g
(,)Tj
0.83 0.64 0.02 0 k
0.9241 0 Td
(4)Tj
0 g
(,)Tj
0.83 0.64 0.02 0 k
0.9184 0 Td
(5)Tj
0 g
(].)Tj
1.2642 0 Td
(Atlantic)Tj
3.4186 0 Td
(sturgeon)Tj
3.7247 0 Td
(\(AS\))Tj
2.0579 0 Td
(were)Tj
2.1203 0 Td
(very)Tj
1.9276 0 Td
(abundant)Tj
4.0422 0 Td
(during)Tj
2.931 0 Td
(European)Tj
4.1158 0 Td
(establishment)Tj
5.7373 0 Td
(of)Tj
1.0148 0 Td
(North)Tj
-35.1153 -1.2983 Td
(America)Tj
3.6396 0 Td
(but)Tj
1.5534 0 Td
(were)Tj
2.126 0 Td
(heavily)Tj
3.0273 0 Td
(exploited)Tj
3.8948 0 Td
(during)Tj
2.9254 0 Td
(the)Tj
1.474 0 Td
(late)Tj
1.6214 0 Td
(1800s,)Tj
2.7099 0 Td
(particularly)Tj
4.8075 0 Td
(for)Tj
1.389 0 Td
(flesh)Tj
2.0636 0 Td
(and)Tj
1.7348 0 Td
(roe)Tj
1.5193 0 Td
([)Tj
0.83 0.64 0.02 0 k
(1)Tj
0 g
(,)Tj
0.83 0.64 0.02 0 k
1.2586 0 Td
(5)Tj
0 g
(,)Tj
0.83 0.64 0.02 0 k
0.9241 0 Td
(6)Tj
0 g
(,)Tj
0.83 0.64 0.02 0 k
-36.6687 -1.3039 Td
(7)Tj
0 g
(,)Tj
0.83 0.64 0.02 0 k
0.9184 0 Td
(8)Tj
0 g
(,)Tj
0.83 0.64 0.02 0 k
0.9184 0 Td
(9)Tj
0 g
(,)Tj
0.83 0.64 0.02 0 k
0.9127 0 Td
(10)Tj
0 g
(].)Tj
1.7348 0 Td
(By)Tj
1.2529 0 Td
(the)Tj
1.4684 0 Td
(end)Tj
1.7121 0 Td
(of)Tj
1.0148 0 Td
(the)Tj
1.4627 0 Td
(19)Tj
ET
Q
q
1 j
1 J
0 w
7.4999 0 0 6.6659 323.4897 641.537 cm
BT
/F6 1 Tf
1 TL
-0.0053 Tc
0 0 Td
(th)Tj
ET
Q
q
1 j
1 J
0 w
10 0 0 10 331.8236 637.0582 cm
BT
/F6 1 Tf
1 TL
-0.004 Tc
0 0 Td
(century)Tj
3.2654 0 Td
(most)Tj
2.2054 0 Td
(of)Tj
1.0091 0 Td
(the)Tj
1.4683 0 Td
(inexhaustible)Tj
6.2986 0 Td
(AS)Tj
1.3719 0 Td
(populations)Tj
4.9493 0 Td
(were)Tj
2.1203 0 Td
(dec-)Tj
-35.8693 -1.2982 Td
(imated)Tj
2.9877 0 Td
(and)Tj
1.7292 0 Td
(most)Tj
2.211 0 Td
(fisheries)Tj
3.5092 0 Td
(collapsed)Tj
3.906 0 Td
([)Tj
0.83 0.64 0.02 0 k
(1)Tj
0 g
(,)Tj
0.83 0.64 0.02 0 k
1.2586 0 Td
(5)Tj
0 g
(,)Tj
0.83 0.64 0.02 0 k
0.9241 0 Td
(6)Tj
0 g
(,)Tj
0.83 0.64 0.02 0 k
0.9184 0 Td
(10)Tj
0 g
(];)Tj
1.7348 0 Td
(however,)Tj
3.8381 0 Td
(these)Tj
2.2564 0 Td
(populations)Tj
4.9492 0 Td
(still)Tj
1.6384 0 Td
(experienced)Tj
-31.8611 -1.2983 Td
(commercial)Tj
4.955 0 Td
(fishing)Tj
2.9423 0 Td
(pressure)Tj
3.5773 0 Td
(until)Tj
2.1089 0 Td
(the)Tj
1.4683 0 Td
(later)Tj
1.9956 0 Td
(part)Tj
1.8425 0 Td
(of)Tj
1.0148 0 Td
(the)Tj
1.4683 0 Td
(20)Tj
ET
Q
q
1 j
1 J
0 w
7.4999 0 0 6.6659 423.2692 615.5149 cm
BT
/F6 1 Tf
1 TL
-0.0053 Tc
0 0 Td
(th)Tj
ET
Q
q
1 j
1 J
0 w
10 0 0 10 431.6031 611.0929 cm
BT
/F6 1 Tf
1 TL
-0.004 Tc
0 0 Td
(century)Tj
3.2711 0 Td
(when)Tj
2.4038 0 Td
(a)Tj
0.6519 0 Td
(moratorium)Tj
5.1421 0 Td
(was)Tj
-34.6278 -1.3039 Td
(imposed)Tj
3.6397 0 Td
(by)Tj
1.1849 0 Td
(the)Tj
1.4683 0 Td
(Atlantic)Tj
3.4186 0 Td
(States)Tj
2.5115 0 Td
(Marine)Tj
3.1577 0 Td
(Fisheries)Tj
3.7531 0 Td
(Commission)Tj
5.3403 0 Td
([)Tj
0.83 0.64 0.02 0 k
(11)Tj
0 g
(].)Tj
2.075 0 Td
(Unfortunately,)Tj
6.1284 0 Td
(as)Tj
1.0148 0 Td
(a)Tj
0.652 0 Td
(result)Tj
2.4491 0 Td
(of)Tj
-36.7934 -1.2982 Td
(AS's)Tj
1.9559 0 Td
(long)Tj
1.9843 0 Td
(generation)Tj
4.507 0 Td
(time)Tj
2.0296 0 Td
(in)Tj
1.0375 0 Td
(combination)Tj
5.3461 0 Td
(with)Tj
1.9956 0 Td
(continued)Tj
4.2916 0 Td
(habitat)Tj
2.9876 0 Td
(impairment)Tj
4.989 0 Td
(such)Tj
2.0579 0 Td
(as)Tj
1.0148 0 Td
(dams)Tj
-34.1969 -1.2983 Td
(and)Tj
1.7292 0 Td
(sedimentation,)Tj
6.1681 0 Td
(AS)Tj
1.3833 0 Td
(populations)Tj
4.9493 0 Td
(have)Tj
2.0579 0 Td
(been)Tj
2.126 0 Td
(very)Tj
1.9219 0 Td
(slow)Tj
2.0182 0 Td
(to)Tj
1.0261 0 Td
(recover)Tj
3.1974 0 Td
([)Tj
0.83 0.64 0.02 0 k
(4)Tj
0 g
(,)Tj
0.83 0.64 0.02 0 k
1.2586 0 Td
(12)Tj
0 g
(,)Tj
0.83 0.64 0.02 0 k
1.3946 0 Td
(13)Tj
0 g
(])Tj
1.5137 0 Td
(which)Tj
2.6476 0 Td
(resulted)Tj
-33.3919 -1.2983 Td
(in)Tj
1.0318 0 Td
(all)Tj
1.1509 0 Td
(US)Tj
1.4287 0 Td
(distinct)Tj
3.2144 0 Td
(population)Tj
4.5865 0 Td
(segments)Tj
3.9231 0 Td
(being)Tj
2.4321 0 Td
(listed)Tj
2.3414 0 Td
(as)Tj
1.0148 0 Td
(either)Tj
2.534 0 Td
(threatened)Tj
4.4958 0 Td
(or)Tj
1.0998 0 Td
(endangered)Tj
4.9096 0 Td
(by)Tj
1.1848 0 Td
(the)Tj
-35.3477 -1.3039 Td
(National)Tj
3.6907 0 Td
(Oceanic)Tj
3.4866 0 Td
(and)Tj
1.7291 0 Td
(Atmospheric)Tj
5.4198 0 Td
(Administration)Tj
6.4233 0 Td
(in)Tj
1.0318 0 Td
(2012)Tj
2.1259 0 Td
([)Tj
0.83 0.64 0.02 0 k
(14)Tj
0 g
(,)Tj
0.83 0.64 0.02 0 k
1.7348 0 Td
(15)Tj
0 g
(].)Tj
-24.4457 -1.2982 Td
(Previously,)Tj
4.6204 0 Td
(it)Tj
0.7824 0 Td
(was)Tj
1.7007 0 Td
(commonly)Tj
4.5468 0 Td
(accepted)Tj
3.6907 0 Td
(that)Tj
1.7801 0 Td
(AS)Tj
1.3776 0 Td
(only)Tj
1.9786 0 Td
(spawned)Tj
3.7133 0 Td
(during)Tj
2.931 0 Td
(the)Tj
1.4683 0 Td
(spring)Tj
2.7496 0 Td
(along)Tj
2.4208 0 Td
(with)Tj
-34.9566 -1.2983 Td
(most)Tj
2.2054 0 Td
(of)Tj
1.0148 0 Td
(the)Tj
1.4683 0 Td
(other)Tj
2.3471 0 Td
(anadromous)Tj
5.2611 0 Td
(fish)Tj
1.661 0 Td
(on)Tj
1.2756 0 Td
(the)Tj
1.4684 0 Td
(east)Tj
1.7404 0 Td
(coast)Tj
2.2394 0 Td
(of)Tj
1.0148 0 Td
(the)Tj
1.4739 0 Td
(US)Tj
1.4286 0 Td
([)Tj
0.83 0.64 0.02 0 k
(1)Tj
0 g
(,)Tj
0.83 0.64 0.02 0 k
1.2643 0 Td
(4)Tj
0 g
(].)Tj
1.2586 0 Td
(However,)Tj
4.0762 0 Td
(relatively)Tj
-31.1979 -1.3039 Td
(recently)Tj
3.4073 0 Td
(researchers)Tj
4.7111 0 Td
(have)Tj
2.0636 0 Td
(shown)Tj
2.846 0 Td
(via)Tj
1.3776 0 Td
(several)Tj
2.931 0 Td
(lines)Tj
2.0693 0 Td
(of)Tj
1.0148 0 Td
(telemetry)Tj
3.9797 0 Td
(evidence)Tj
3.7134 0 Td
(that)Tj
1.7745 0 Td
(some)Tj
2.33 0 Td
(rivers)Tj
2.4718 0 Td
(likely)Tj
-34.6901 -1.2983 Td
(have)Tj
2.058 0 Td
(dual-spawning)Tj
6.1341 0 Td
(groups)Tj
2.9707 0 Td
(of)Tj
1.0148 0 Td
(AS,)Tj
1.6044 0 Td
(one)Tj
1.6951 0 Td
(group)Tj
2.6079 0 Td
(spawning)Tj
4.0478 0 Td
(in)Tj
1.0374 0 Td
(the)Tj
1.4683 0 Td
(spring)Tj
2.7496 0 Td
(and)Tj
1.7291 0 Td
(another)Tj
3.3336 0 Td
(group)Tj
-32.4508 -1.2983 Td
(spawning)Tj
4.0422 0 Td
(in)Tj
1.0375 0 Td
(the)Tj
1.4683 0 Td
(fall)Tj
1.4344 0 Td
([)Tj
0.83 0.64 0.02 0 k
(10)Tj
0 g
(,)Tj
0.83 0.64 0.02 0 k
1.7404 0 Td
(16)Tj
0 g
(,)Tj
0.83 0.64 0.02 0 k
1.3947 0 Td
(17)Tj
0 g
(,)Tj
0.83 0.64 0.02 0 k
1.3946 0 Td
(18)Tj
0 g
(].)Tj
1.7404 0 Td
(Additionally,)Tj
5.4539 0 Td
(river-resident)Tj
5.6805 0 Td
(juvenile)Tj
3.3675 0 Td
(AS)Tj
1.3776 0 Td
(\()Tj
/F12 1 Tf
0 Tc
(<)Tj
/F6 1 Tf
-0.004 Tc
(530)Tj
2.761 0 Td
(mm)Tj
1.8595 0 Td
(TL\))Tj
-34.7525 -1.3039 Td
(from)Tj
2.2054 0 Td
(the)Tj
1.4683 0 Td
(Edisto)Tj
2.7496 0 Td
(River,)Tj
2.5852 0 Td
(SC)Tj
1.3606 0 Td
(were)Tj
2.1203 0 Td
(assigned)Tj
3.6057 0 Td
(to)Tj
1.0261 0 Td
(either)Tj
2.5342 0 Td
(fall)Tj
1.4343 0 Td
(or)Tj
1.0998 0 Td
(spring)Tj
2.7495 0 Td
(spawning)Tj
4.0422 0 Td
(events)Tj
2.7326 0 Td
(based)Tj
2.4718 0 Td
(on)Tj
1.2755 0 Td
(total)Tj
-35.4611 -1.2982 Td
(length)Tj
2.727 0 Td
(at)Tj
0.9524 0 Td
(day)Tj
1.6384 0 Td
(of)Tj
1.0148 0 Td
(year)Tj
1.9049 0 Td
(over)Tj
1.9729 0 Td
(a)Tj
0.6576 0 Td
(15)Tj
1.1679 0 Td
(year)Tj
1.9048 0 Td
(period)Tj
2.8347 0 Td
(and)Tj
1.7291 0 Td
(subsequently)Tj
5.4084 0 Td
(determined)Tj
4.8472 0 Td
(to)Tj
1.0261 0 Td
(be)Tj
1.1509 0 Td
(genetically)Tj
4.4503 0 Td
(dis-)Tj
-35.3874 -1.2983 Td
(tinct)Tj
2.0523 0 Td
(with)Tj
2.0013 0 Td
(little)Tj
1.9955 0 Td
(admixture)Tj
4.3597 0 Td
(between)Tj
3.5263 0 Td
(the)Tj
1.4683 0 Td
(spawning)Tj
4.0479 0 Td
(groups)Tj
2.9706 0 Td
([)Tj
0.83 0.64 0.02 0 k
(19)Tj
0 g
(].)Tj
2.0749 0 Td
(Currently)Tj
4.1272 0 Td
(dual-spawning)Tj
6.1285 0 Td
(in)Tj
1.0375 0 Td
(the)Tj
-35.79 -1.2983 Td
(James)Tj
2.5796 0 Td
(River,)Tj
2.5795 0 Td
(VA)Tj
1.593 0 Td
(has)Tj
1.5364 0 Td
(been)Tj
2.1203 0 Td
(documented)Tj
5.2441 0 Td
(based)Tj
2.4661 0 Td
(on)Tj
1.2699 0 Td
(collections)Tj
4.4673 0 Td
(of)Tj
1.0091 0 Td
(pre-)Tj
1.8765 0 Td
(and)Tj
1.7291 0 Td
(post-spawn)Tj
4.7906 0 Td
(AS)Tj
1.3776 0 Td
(accom-)Tj
-34.6391 -1.3039 Td
(panied)Tj
2.9367 0 Td
(by)Tj
1.1792 0 Td
(telemetry)Tj
3.9798 0 Td
(data)Tj
1.9162 0 Td
([)Tj
0.83 0.64 0.02 0 k
(10)Tj
0 g
(,)Tj
0.83 0.64 0.02 0 k
1.7348 0 Td
(18)Tj
0 g
(].)Tj
1.7405 0 Td
(Due)Tj
1.9049 0 Td
(to)Tj
1.0261 0 Td
(the)Tj
1.4683 0 Td
(lack)Tj
1.8199 0 Td
(of)Tj
1.0148 0 Td
(juvenile)Tj
3.3674 0 Td
(AS)Tj
1.3833 0 Td
(captures)Tj
3.5773 0 Td
(in)Tj
1.0318 0 Td
(the)Tj
1.474 0 Td
(James)Tj
2.5852 0 Td
(River,)Tj
-34.1402 -1.2982 Td
(genetics)Tj
3.4299 0 Td
(from)Tj
2.2054 0 Td
(the)Tj
1.4683 0 Td
(two)Tj
1.7121 0 Td
(spawning)Tj
4.0422 0 Td
(populations)Tj
4.9549 0 Td
(cannot)Tj
2.9764 0 Td
(be)Tj
1.1509 0 Td
(compared)Tj
4.2461 0 Td
(in)Tj
1.0375 0 Td
(the)Tj
1.4683 0 Td
(same)Tj
2.2564 0 Td
(river-resident)Tj
-30.9484 -1.2983 Td
(juvenile)Tj
3.3562 0 Td
(design)Tj
2.8063 0 Td
(similar)Tj
2.9764 0 Td
(to)Tj
1.0148 0 Td
(the)Tj
1.4626 0 Td
(Edisto)Tj
2.744 0 Td
(River)Tj
2.347 0 Td
(study;)Tj
2.6079 0 Td
(however,)Tj
3.8267 0 Td
(adult)Tj
2.245 0 Td
(samples)Tj
3.3732 0 Td
(are)Tj
1.44 0 Td
(available)Tj
3.6623 0 Td
(with)Tj
1.9842 0 Td
(sub-)Tj
-35.8466 -1.3039 Td
(stantial)Tj
3.1068 0 Td
(collection)Tj
4.1102 0 Td
(information)Tj
5.074 0 Td
(to)Tj
1.0261 0 Td
(attempt)Tj
3.3109 0 Td
(to)Tj
1.0261 0 Td
(genetically)Tj
4.4504 0 Td
(address)Tj
3.2257 0 Td
(these)Tj
2.2563 0 Td
(putative)Tj
3.4413 0 Td
(spawning)Tj
-31.0278 -1.2983 Td
(groups.)Tj
3.1918 0 Td
(As)Tj
1.2643 0 Td
(such,)Tj
2.2904 0 Td
(the)Tj
1.4683 0 Td
(main)Tj
2.2904 0 Td
(objective)Tj
3.7814 0 Td
(of)Tj
1.0148 0 Td
(our)Tj
1.6214 0 Td
(study)Tj
2.3924 0 Td
(was)Tj
1.7008 0 Td
(to)Tj
1.0204 0 Td
(determine)Tj
4.3256 0 Td
(if)Tj
0.7767 0 Td
(genetic)Tj
3.067 0 Td
(differences)Tj
4.5638 0 Td
(were)Tj
-34.7695 -1.2983 Td
(present)Tj
3.1635 0 Td
(between)Tj
3.5263 0 Td
(spring)Tj
2.7496 0 Td
(and)Tj
1.7291 0 Td
(fall)Tj
1.4286 0 Td
(run)Tj
1.6668 0 Td
(AS)Tj
1.3833 0 Td
(by)Tj
1.1792 0 Td
(sampling)Tj
3.8778 0 Td
(the)Tj
1.4683 0 Td
(adult)Tj
2.2506 0 Td
(populations)Tj
4.9493 0 Td
(during)Tj
2.9253 0 Td
(the)Tj
1.474 0 Td
(spawning)Tj
-33.7717 -1.3039 Td
(seasons.)Tj
3.4413 0 Td
(Secondary)Tj
4.3823 0 Td
(objectives)Tj
4.1442 0 Td
(included)Tj
3.7134 0 Td
(determining)Tj
5.1817 0 Td
(the)Tj
1.4683 0 Td
(genetic)Tj
3.067 0 Td
(health)Tj
2.6816 0 Td
(of)Tj
1.0148 0 Td
(the)Tj
1.4683 0 Td
(population\(s\))Tj
-30.5629 -1.2982 Td
(and)Tj
1.7292 0 Td
(evaluating)Tj
4.3199 0 Td
(if)Tj
0.7767 0 Td
(the)Tj
1.4684 0 Td
(use)Tj
1.5307 0 Td
(of)Tj
1.0148 0 Td
(appropriate)Tj
4.8812 0 Td
(sample)Tj
3.0217 0 Td
(selection)Tj
3.7134 0 Td
(criteria)Tj
3.0783 0 Td
(with)Tj
1.9955 0 Td
(adult)Tj
2.2564 0 Td
(AS)Tj
1.3776 0 Td
(can)Tj
1.6271 0 Td
(be)Tj
1.1509 0 Td
(used)Tj
2.0579 0 Td
(to)Tj
-35.9997 -1.2983 Td
(identify)Tj
3.2939 0 Td
(dual-spawning)Tj
6.1341 0 Td
(group)Tj
2.6079 0 Td
(genetic)Tj
3.067 0 Td
(signatures.)Tj
ET
Q
q
1 j
1 J
0 w
11.9999 0 0 11.9999 200.0125 282.0472 cm
BT
/F2 1 Tf
1 TL
-0.0033 Tc
0 0 Td
(Study)Tj
2.9811 0 Td
(area)Tj
ET
Q
q
1 j
1 J
0 w
10 0 0 10 200.0125 265.0393 cm
BT
/F6 1 Tf
1 TL
-0.004 Tc
0 0 Td
(The)Tj
1.7801 0 Td
(James)Tj
2.5908 0 Td
(River,)Tj
2.5796 0 Td
(river)Tj
2.1089 0 Td
(mouth)Tj
2.9027 0 Td
(located)Tj
3.0784 0 Td
(at)Tj
0.9524 0 Td
(N)Tj
0.9581 0 Td
(36.98889,)Tj
3.9968 0 Td
(W)Tj
1.1849 0 Td
(76.30365,)Tj
4.0025 0 Td
(is)Tj
0.8504 0 Td
(the)Tj
1.4683 0 Td
(southern-most)Tj
6.1228 0 Td
(major)Tj
-34.5767 -1.2982 Td
(tributary)Tj
3.7473 0 Td
(of)Tj
1.0148 0 Td
(the)Tj
1.4684 0 Td
(Chesapeake)Tj
4.9265 0 Td
(Bay,)Tj
1.9163 0 Td
(is)Tj
0.8447 0 Td
(696)Tj
1.6497 0 Td
(km)Tj
1.5364 0 Td
(long)Tj
1.9842 0 Td
(and)Tj
1.7292 0 Td
(drains)Tj
2.7325 0 Td
(26,164)Tj
2.8233 0 Td
(km)Tj
ET
Q
q
1 j
1 J
0 w
7.4999 0 0 6.6659 476.9007 256.4787 cm
BT
/F6 1 Tf
1 TL
-0.0053 Tc
0 0 Td
(2)Tj
ET
Q
q
1 j
1 J
0 w
10 0 0 10 480.4724 252.0566 cm
BT
/F6 1 Tf
1 TL
-0.004 Tc
0 0 Td
(.)Tj
0.4422 0 Td
(The)Tj
1.7858 0 Td
(tidal)Tj
1.9899 0 Td
(portion)Tj
3.2371 0 Td
(of)Tj
-35.5008 -1.3039 Td
(the)Tj
1.4684 0 Td
(river)Tj
2.1089 0 Td
(extends)Tj
3.2655 0 Td
(up)Tj
1.2643 0 Td
(to)Tj
1.0261 0 Td
(Richmond,)Tj
4.6885 0 Td
(VA)Tj
1.5987 0 Td
(located)Tj
3.0841 0 Td
(at)Tj
0.9468 0 Td
(rkm)Tj
1.9105 0 Td
(155.)Tj
1.8709 0 Td
(The)Tj
1.7858 0 Td
(river)Tj
2.1033 0 Td
(width)Tj
2.5283 0 Td
(varies)Tj
2.5285 0 Td
(between)Tj
3.532 0 Td
(0.7)Tj
-35.7106 -1.2982 Td
(and)Tj
1.7292 0 Td
(7.1km)Tj
2.7099 0 Td
(up)Tj
1.2699 0 Td
(to)Tj
1.0204 0 Td
(rkm)Tj
1.9106 0 Td
(120)Tj
1.6441 0 Td
(and)Tj
1.7348 0 Td
(then)Tj
2.0182 0 Td
(narrows)Tj
3.498 0 Td
(substantially)Tj
5.1987 0 Td
(\(0.10.4)Tj
3.4355 0 Td
(km\).)Tj
2.0976 0 Td
(The)Tj
1.7858 0 Td
(federal)Tj
2.931 0 Td
(navigation)Tj
-32.9837 -1.2983 Td
(channel)Tj
3.3732 0 Td
(maintained)Tj
4.7849 0 Td
(by)Tj
1.1849 0 Td
(the)Tj
1.4683 0 Td
(US)Tj
1.4287 0 Td
(Army)Tj
2.5511 0 Td
(Corps)Tj
2.6419 0 Td
(of)Tj
1.0148 0 Td
(Engineers)Tj
4.1952 0 Td
(runs)Tj
2.0296 0 Td
(from)Tj
2.2054 0 Td
(the)Tj
1.4739 0 Td
(river)Tj
2.1033 0 Td
(mouth)Tj
2.9026 0 Td
(at)Tj
0.9525 0 Td
(Hamp-)Tj
-34.3103 -1.3039 Td
(ton)Tj
1.5704 0 Td
(Roads,)Tj
2.897 0 Td
(VA)Tj
1.5987 0 Td
(to)Tj
1.0262 0 Td
(rkm)Tj
1.9105 0 Td
(150)Tj
1.6441 0 Td
(and)Tj
1.7291 0 Td
(is)Tj
0.8504 0 Td
(maintained)Tj
4.7905 0 Td
(to)Tj
1.0205 0 Td
(a)Tj
0.6576 0 Td
(minimum)Tj
4.286 0 Td
(depth)Tj
2.5171 0 Td
(of)Tj
1.0148 0 Td
(7.6)Tj
1.3946 0 Td
(m)Tj
1.0431 0 Td
(and)Tj
1.7291 0 Td
(a)Tj
0.652 0 Td
(minimum)Tj
-32.3317 -1.2983 Td
(width)Tj
2.5229 0 Td
(of)Tj
1.0148 0 Td
(91.4)Tj
1.8708 0 Td
(m.)Tj
1.2699 0 Td
(AS)Tj
1.3777 0 Td
(in)Tj
1.0374 0 Td
(the)Tj
1.4684 0 Td
(James)Tj
2.5851 0 Td
(River)Tj
2.3585 0 Td
(were)Tj
2.1259 0 Td
(exploited)Tj
3.8948 0 Td
(by)Tj
1.1792 0 Td
(Europeans)Tj
4.4787 0 Td
(since)Tj
2.2449 0 Td
(the)Tj
1.4684 0 Td
(founding)Tj
3.9004 0 Td
(of)Tj
-34.7978 -1.2982 Td
(Jamestown)Tj
4.6262 0 Td
(Colony)Tj
3.1464 0 Td
(in)Tj
1.0318 0 Td
(1607)Tj
2.126 0 Td
([)Tj
0.83 0.64 0.02 0 k
(20)Tj
0 g
(])Tj
1.8481 0 Td
(and)Tj
1.7348 0 Td
(harvest)Tj
3.0898 0 Td
(continued)Tj
4.2916 0 Td
(in)Tj
1.0318 0 Td
(some)Tj
2.3357 0 Td
(capacity)Tj
3.4695 0 Td
(until)Tj
2.109 0 Td
(1974)Tj
2.1203 0 Td
(when)Tj
2.4038 0 Td
(the)Tj
-35.3648 -1.2983 Td
(Virginia)Tj
3.532 0 Td
(Marine)Tj
3.1578 0 Td
(Resources)Tj
4.2406 0 Td
(Commission)Tj
5.3404 0 Td
(implemented)Tj
5.5389 0 Td
(a)Tj
0.6519 0 Td
(state-wide)Tj
4.286 0 Td
(ban)Tj
1.712 0 Td
(on)Tj
1.2756 0 Td
(all)Tj
1.1452 0 Td
(AS)Tj
1.3776 0 Td
(harvest)Tj
3.0954 0 Td
([)Tj
0.83 0.64 0.02 0 k
(21)Tj
0 g
(].)Tj
ET
Q
q
1 j
1 J
0 w
11.9999 0 0 11.9999 200.0125 131.0173 cm
BT
/F2 1 Tf
1 TL
-0.0033 Tc
0 0 Td
(Materials)Tj
4.5071 0 Td
(and)Tj
1.989 0 Td
(methods)Tj
/F0 1 Tf
-6.4961 -1.5023 Td
(Genetic)Tj
3.6283 0 Td
(sample)Tj
3.4064 0 Td
(collection)Tj
ET
Q
q
1 j
1 J
0 w
10 0 0 10 200.0125 95.9811 cm
BT
/F6 1 Tf
1 TL
-0.004 Tc
0 0 Td
(Since)Tj
2.3584 0 Td
(October)Tj
3.4979 0 Td
(2007,)Tj
2.3471 0 Td
(gill)Tj
1.4456 0 Td
(net)Tj
1.491 0 Td
(sampling)Tj
3.8778 0 Td
(targeting)Tj
3.7927 0 Td
(adult)Tj
2.2507 0 Td
(AS)Tj
1.3777 0 Td
(has)Tj
1.5477 0 Td
(been)Tj
2.1203 0 Td
(conducted)Tj
4.4277 0 Td
(in)Tj
1.0374 0 Td
(the)Tj
1.4684 0 Td
(James)Tj
-33.0404 -1.2982 Td
(River)Tj
2.3584 0 Td
(during)Tj
2.9253 0 Td
(the)Tj
1.4683 0 Td
(spring)Tj
2.7553 0 Td
(and)Tj
1.7291 0 Td
(fall)Tj
1.4343 0 Td
(season.)Tj
3.0784 0 Td
(Sampling)Tj
3.9969 0 Td
(protocols)Tj
3.9571 0 Td
(and)Tj
1.7291 0 Td
(tagging)Tj
3.1635 0 Td
(techniques)Tj
4.5297 0 Td
(are)Tj
ET
Q
q
1 j
1 J
0 w
576 737.1 m
36 737.1 l
36 737.6 l
576 737.6 l
f*
36 741.2598 107.1496 23.6976 re
W* n
q
107.0929 0 0 23.6409 36 741.3165 cm
q
/I0 Do
Q
Q
Q
q
0 0 612 792 re
W* n
1 j
1 J
0 w
7.9999 0 0 7.9999 317.3669 745.7952 cm
BT
/F0 1 Tf
1 TL
-0.005 Tc
0 0 Td
(Genetic)Tj
3.6071 0 Td
(differentiation)Tj
6.1087 0 Td
(of)Tj
1.0417 0 Td
(male)Tj
2.3457 0 Td
(Atlantic)Tj
3.4441 0 Td
(sturgeon)Tj
4.0394 0 Td
(in)Tj
0.978 0 Td
(the)Tj
1.5874 0 Td
(James)Tj
3.1252 0 Td
(River,)Tj
2.7851 0 Td
(Virginia)Tj
ET
Q
q
1 j
1 J
0 w
36 48.0002 m
576 48.0002 l
576 47.5002 l
36 47.5002 l
f*
0.83 0.64 0.02 0 k
81.5811 34.9228 m
237.8835 34.9228 l
h
f*
0 g
7.9999 0 0 7.9999 36 36 cm
BT
/F0 1 Tf
1 TL
-0.005 Tc
0 0 Td
(PLOS)Tj
2.8559 0 Td
(ONE)Tj
2.3669 0 Td
(|)Tj
0.83 0.64 0.02 0 k
0.4748 0 Td
(https://doi.or)Tj
5.4142 0 Td
(g/10.137)Tj
3.8198 0 Td
(1/journal.po)Tj
5.1236 0 Td
(ne.01796)Tj
4.089 0 Td
(61)Tj
0 g
2.0906 0 Td
(July)Tj
1.9701 0 Td
(7,)Tj
1.0417 0 Td
(2017)Tj
36.4397 0 Td
(2)Tj
0.7654 0 Td
(/)Tj
0.496 0 Td
(8)Tj
/F8 1 Tf
-0.025 Tc
-66.9477 84.0904 Td
(Competing)Tj
4.3795 0 Td
(interests)Tj
3.3307 0 Td
(:)Tj
/F9 1 Tf
0.4819 0 Td
(The)Tj
1.5875 0 Td
(authors)Tj
3.0401 0 Td
(have)Tj
1.963 0 Td
(declared)Tj
-14.7827 -1.3748 Td
(that)Tj
1.6228 0 Td
(no)Tj
1.1693 0 Td
(competing)Tj
4.1599 0 Td
(interests)Tj
ET
endstream
endobj
52 0 obj
<>stream
endstream
endobj
53 0 obj
<>stream
endstream
endobj
54 0 obj
<>stream
endstream
endobj
55 0 obj
<>stream
endstream
endobj
56 0 obj
<>stream
endstream
endobj
57 0 obj
<>stream
endstream
endobj
58 0 obj
<>stream
endstream
endobj
59 0 obj
<>stream
endstream
endobj
60 0 obj
<>stream
endstream
endobj
61 0 obj
<>stream
endstream
endobj
62 0 obj
<>stream
endstream
endobj
63 0 obj
<>stream
endstream
endobj
64 0 obj
<>stream
endstream
endobj
65 0 obj
<>stream
endstream
endobj
66 0 obj
<>stream
endstream
endobj
67 0 obj
<>stream
endstream
endobj
68 0 obj
<>stream
endstream
endobj
69 0 obj
<>stream
endstream
endobj
70 0 obj
<>stream
endstream
endobj
71 0 obj
<>stream
endstream
endobj
72 0 obj
<>stream
endstream
endobj
73 0 obj
<>stream
endstream
endobj
74 0 obj
<>stream
endstream
endobj
75 0 obj
<>stream
endstream
endobj
76 0 obj
<>stream
BT
10.3394 82.7156 Td
(exist.)Tj
ET
Q
endstream
endobj
77 0 obj
<>/XObject<>>>/CropBox[0 0 612 792]/MediaBox[0 0 612 792]/Parent 13 0 R/Annots 78 0 R/Contents 79 0 R/TrimBox[0 0 612 792]>>
endobj
78 0 obj
[80 0 R 81 0 R 82 0 R 83 0 R 84 0 R 85 0 R 86 0 R 87 0 R 88 0 R 89 0 R 90 0 R 91 0 R 92 0 R 93 0 R 94 0 R 95 0 R]
endobj
80 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref010)>>
endobj
81 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref018)>>
endobj
82 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref022)>>
endobj
83 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref010)>>
endobj
84 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref022)>>
endobj
85 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref018)>>
endobj
86 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref019)>>
endobj
87 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref023)>>
endobj
88 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref024)>>
endobj
89 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref025)>>
endobj
90 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref026)>>
endobj
91 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref027)>>
endobj
92 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref028)>>
endobj
93 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref029)>>
endobj
94 0 obj
<>/Border[0 0 0]/Dest(Rpone.0179661.ref030)>>
endobj
95 0 obj
<>/Border[0 0 0]/A 96 0 R>>
endobj
96 0 obj
<>
endobj
79 0 obj
[97 0 R 98 0 R 99 0 R 100 0 R 101 0 R 102 0 R 103 0 R 104 0 R 105 0 R 106 0 R 107 0 R 108 0 R 109 0 R 110 0 R]
endobj
97 0 obj
<>stream
q
0.83 0.64 0.02 0 k
339.5339 706.337 m
348.9449 706.337 l
h
f*
353.2535 706.337 m
362.6079 706.337 l
h
f*
366.9732 706.337 m
376.3276 706.337 l
h
f*
535.8614 589.3228 m
545.3858 589.3228 l
h
f*
549.8079 589.3228 m
559.3323 589.3228 l
h
f*
376.2142 459.326 m
385.7386 459.326 l
h
f*
409.0961 393.9024 m
418.6205 393.9024 l
h
f*
408.1323 250.9228 m
417.6567 250.9228 l
h
f*
304.6677 185.8961 m
314.1921 185.8961 l
h
f*
313.5685 172.9134 m
323.0362 172.9134 l
h
f*
265.6063 133.9087 m
275.1307 133.9087 l
h
f*
279.5528 133.9087 m
289.0772 133.9087 l
h
f*
361.7575 120.926 m
371.2819 120.926 l
h
f*
375.7039 120.926 m
385.2283 120.926 l
h
f*
500.2583 81.9213 m
509.726 81.9213 l
h
f*
0 g
1 j
1 J
0 w
10 0 0 10 200.0125 707.4141 cm
BT
/F6 1 Tf
1 TL
-0.011 Tc
0 0 Td
(described)Tj
3.9798 0 Td
(in)Tj
1.0204 0 Td
(previous)Tj
3.583 0 Td
(publications)Tj
5.04 0 Td
([)Tj
0.83 0.64 0.02 0 k
(10)Tj
0 g
(,)Tj
0.83 0.64 0.02 0 k
1.7007 0 Td
(18)Tj
0 g
(,)Tj
0.83 0.64 0.02 0 k
1.372 0 Td
(22)Tj
0 g
(].)Tj
1.7008 0 Td
(All)Tj
1.3719 0 Td
(sampling)Tj
3.8155 0 Td
(and)Tj
1.7007 0 Td
(fish)Tj
1.6328 0 Td
(workup)Tj
3.2768 0 Td
(was)Tj
1.6724 0 Td
(conducted)Tj
4.3597 0 Td
(fol-)Tj
-36.2265 -1.3039 Td
(lowing)Tj
2.8969 0 Td
(protocols)Tj
3.8948 0 Td
(set)Tj
1.2813 0 Td
(by)Tj
1.1678 0 Td
(Virginia)Tj
3.4753 0 Td
(Commonwealth)Tj
6.616 0 Td
(University's)Tj
4.8982 0 Td
(IACUC)Tj
3.2712 0 Td
(#AD20127)Tj
4.439 0 Td
(and)Tj
1.7065 0 Td
(National)Tj
-33.647 -1.2982 Td
(Oceanic)Tj
3.4355 0 Td
(and)Tj
1.7121 0 Td
(Atmospheric)Tj
5.3405 0 Td
(Administration)Tj
6.3212 0 Td
(endangered)Tj
4.8472 0 Td
(species)Tj
2.948 0 Td
(permit)Tj
2.8743 0 Td
(#16547.)Tj
3.2485 0 Td
(From)Tj
2.4208 0 Td
(2007)Tj
2.0919 0 Td
(to)Tj
-35.24 -1.2983 Td
(2016,)Tj
2.313 0 Td
(a)Tj
0.6463 0 Td
(total)Tj
1.9672 0 Td
(of)Tj
1.0035 0 Td
(507)Tj
1.6214 0 Td
(unique)Tj
2.982 0 Td
(fall)Tj
1.406 0 Td
(run)Tj
1.6498 0 Td
(AS)Tj
1.3606 0 Td
(were)Tj
2.0976 0 Td
(captured)Tj
3.685 0 Td
(and)Tj
1.7121 0 Td
(40)Tj
1.1566 0 Td
(unique)Tj
2.982 0 Td
(AS)Tj
1.3606 0 Td
(were)Tj
2.0976 0 Td
(captured)Tj
3.6851 0 Td
(during)Tj
-33.7264 -1.2983 Td
(the)Tj
1.4456 0 Td
(spring)Tj
2.7099 0 Td
(run.)Tj
1.8652 0 Td
(Fin)Tj
1.542 0 Td
(clips)Tj
2.0013 0 Td
(were)Tj
2.0919 0 Td
(taken)Tj
2.3868 0 Td
(from)Tj
2.177 0 Td
(most)Tj
2.177 0 Td
(of)Tj
1.0034 0 Td
(the)Tj
1.4514 0 Td
(AS)Tj
1.3606 0 Td
(captured)Tj
3.6907 0 Td
(during)Tj
2.8856 0 Td
(the)Tj
1.4457 0 Td
(study.)Tj
2.5738 0 Td
(On)Tj
1.5024 0 Td
(occa-)Tj
-34.3103 -1.3039 Td
(sion,)Tj
2.0919 0 Td
(fin)Tj
1.3039 0 Td
(clips)Tj
2.0013 0 Td
(for)Tj
1.3606 0 Td
(genetic)Tj
3.0217 0 Td
(samples)Tj
3.3335 0 Td
(were)Tj
2.0977 0 Td
(not)Tj
1.5533 0 Td
(taken)Tj
2.3868 0 Td
(due)Tj
1.6724 0 Td
(to)Tj
1.0148 0 Td
(high)Tj
1.9786 0 Td
(catch)Tj
2.2904 0 Td
(numbers)Tj
3.7303 0 Td
(and)Tj
1.7122 0 Td
(time)Tj
2.0012 0 Td
(con-)Tj
-33.5506 -1.2982 Td
(straints;)Tj
3.3221 0 Td
(however,)Tj
3.7814 0 Td
(unique)Tj
2.9764 0 Td
(passive-integrated-transpon)Tj
10.9757 0 Td
(der)Tj
1.5193 0 Td
(tags)Tj
1.7518 0 Td
(were)Tj
2.0977 0 Td
(placed)Tj
2.7495 0 Td
(in)Tj
1.0205 0 Td
(every)Tj
2.3131 0 Td
(AS)Tj
1.3663 0 Td
(captured.)Tj
-0.004 Tc
-32.6776 -1.2983 Td
(From)Tj
2.4491 0 Td
(the)Tj
1.4683 0 Td
(507)Tj
1.6498 0 Td
(fall)Tj
1.4343 0 Td
(collections,)Tj
4.6941 0 Td
(116)Tj
1.6498 0 Td
(males)Tj
2.5001 0 Td
(were)Tj
2.126 0 Td
(selected)Tj
3.3449 0 Td
(for)Tj
1.3889 0 Td
(genetic)Tj
3.0671 0 Td
(analysis.)Tj
3.5546 0 Td
(The)Tj
1.7802 0 Td
(116)Tj
1.6441 0 Td
(\(2012)Tj
-33.9475 -1.3039 Td
(n)Tj
0.7653 0 Td
(=)Tj
0.7994 0 Td
(70,)Tj
1.4003 0 Td
(2013)Tj
2.1203 0 Td
(n)Tj
0.771 0 Td
(=)Tj
0.7994 0 Td
(46\))Tj
1.508 0 Td
(males)Tj
2.5058 0 Td
(were)Tj
2.126 0 Td
(picked)Tj
2.8743 0 Td
(because)Tj
3.3165 0 Td
(each)Tj
2.0296 0 Td
(had)Tj
1.7121 0 Td
(a)Tj
0.652 0 Td
(telemetry)Tj
3.9798 0 Td
(tag)Tj
1.4173 0 Td
(surgically)Tj
4.0195 0 Td
(implanted.)Tj
-32.7966 -1.2983 Td
(All)Tj
1.3946 0 Td
(fish)Tj
1.6667 0 Td
(were)Tj
2.1203 0 Td
(captured)Tj
3.7418 0 Td
(close)Tj
2.1826 0 Td
(to)Tj
1.0262 0 Td
(spawning)Tj
4.0421 0 Td
(habitat)Tj
2.9877 0 Td
(and)Tj
1.7292 0 Td
(expelled)Tj
3.4979 0 Td
(sperm)Tj
2.7099 0 Td
(during)Tj
2.931 0 Td
(capture)Tj
3.2145 0 Td
([)Tj
0.83 0.64 0.02 0 k
(10)Tj
0 g
(,)Tj
0.83 0.64 0.02 0 k
1.7348 0 Td
(22)Tj
0 g
(].)Tj
-34.9793 -1.2983 Td
(Resulting)Tj
3.9798 0 Td
(real-time)Tj
3.8551 0 Td
(telemetry)Tj
3.9741 0 Td
(data)Tj
1.9162 0 Td
(show)Tj
2.3017 0 Td
(all)Tj
1.1452 0 Td
(but)Tj
1.5534 0 Td
(one)Tj
1.7008 0 Td
(fish)Tj
1.6611 0 Td
(routinely)Tj
3.8664 0 Td
(returned)Tj
3.7077 0 Td
(to)Tj
1.0261 0 Td
(James)Tj
2.5909 0 Td
(River)Tj
-33.2785 -1.3039 Td
(spawning)Tj
4.0421 0 Td
(habitat)Tj
2.9877 0 Td
(during)Tj
2.931 0 Td
(the)Tj
1.4684 0 Td
(fall)Tj
1.4343 0 Td
(spawning)Tj
4.0422 0 Td
(season.)Tj
3.084 0 Td
(The)Tj
1.7802 0 Td
(one)Tj
1.7008 0 Td
(fish)Tj
1.6611 0 Td
(that)Tj
1.7801 0 Td
(did)Tj
1.542 0 Td
(not)Tj
1.5761 0 Td
(routinely)Tj
3.8608 0 Td
(return)Tj
2.7609 0 Td
(to)Tj
-36.6517 -1.2982 Td
(James)Tj
2.5851 0 Td
(River)Tj
2.3584 0 Td
(spawning)Tj
4.0479 0 Td
(habitat)Tj
2.9877 0 Td
(was)Tj
1.6951 0 Td
(captured)Tj
3.7474 0 Td
(in)Tj
1.0318 0 Td
(2012)Tj
2.1259 0 Td
(and)Tj
1.7292 0 Td
(was)Tj
1.7007 0 Td
(detected)Tj
3.566 0 Td
(at)Tj
0.9524 0 Td
(the)Tj
1.4684 0 Td
(mouth)Tj
2.897 0 Td
(of)Tj
1.0148 0 Td
(the)Tj
-33.9078 -1.2983 Td
(James)Tj
2.5851 0 Td
(River)Tj
2.3584 0 Td
(during)Tj
2.931 0 Td
(the)Tj
1.4684 0 Td
(summer)Tj
3.5433 0 Td
(of)Tj
1.0148 0 Td
(2013)Tj
2.1203 0 Td
(and)Tj
1.7291 0 Td
(2014)Tj
2.126 0 Td
(but)Tj
1.5533 0 Td
(did)Tj
1.5421 0 Td
(not)Tj
1.5704 0 Td
(move)Tj
2.4264 0 Td
(upstream)Tj
3.9741 0 Td
(to)Tj
1.0262 0 Td
(spawning)Tj
-31.9689 -1.3039 Td
(grounds.)Tj
3.753 0 Td
(All)Tj
1.3946 0 Td
(of)Tj
1.0148 0 Td
(the)Tj
1.474 0 Td
(2014)Tj
2.1203 0 Td
(\(n)Tj
1.1055 0 Td
(=)Tj
0.8051 0 Td
(2\),)Tj
1.2586 0 Td
(2015)Tj
2.1203 0 Td
(\(n)Tj
1.1111 0 Td
(=)Tj
0.7994 0 Td
(20\),)Tj
1.7348 0 Td
(and)Tj
1.7348 0 Td
(2016)Tj
2.1203 0 Td
(\(n)Tj
1.1055 0 Td
(=)Tj
0.805 0 Td
(16\))Tj
1.508 0 Td
(spring)Tj
2.7496 0 Td
(male)Tj
2.143 0 Td
(collections)Tj
-30.8577 -1.2983 Td
(\(n)Tj
1.1055 0 Td
(=)Tj
0.7993 0 Td
(38\))Tj
1.5137 0 Td
(were)Tj
2.1203 0 Td
(used)Tj
2.0636 0 Td
(for)Tj
1.3833 0 Td
(genetic)Tj
3.0728 0 Td
(analysis)Tj
3.3221 0 Td
(due)Tj
1.7008 0 Td
(to)Tj
1.0262 0 Td
(low)Tj
1.6554 0 Td
(sample)Tj
3.0217 0 Td
(size.)Tj
1.9105 0 Td
(Within)Tj
3.0954 0 Td
(the)Tj
1.4684 0 Td
(spring)Tj
2.7496 0 Td
(sample)Tj
3.0217 0 Td
(set,)Tj
1.5307 0 Td
(18)Tj
-36.561 -1.2982 Td
(of)Tj
1.0147 0 Td
(the)Tj
1.4684 0 Td
(38)Tj
1.1735 0 Td
(spring)Tj
2.7496 0 Td
(samples)Tj
3.3789 0 Td
(have)Tj
2.0636 0 Td
(accompanying)Tj
6.0604 0 Td
(telemetry)Tj
3.9798 0 Td
(data)Tj
1.9163 0 Td
(documenting)Tj
5.5728 0 Td
(repeated)Tj
3.6397 0 Td
(spring)Tj
-33.0177 -1.2983 Td
(spawning)Tj
4.0421 0 Td
(movements.)Tj
5.1024 0 Td
(However,)Tj
4.0762 0 Td
(three)Tj
2.2677 0 Td
(of)Tj
1.0148 0 Td
(the)Tj
1.4683 0 Td
(non-telemetered)Tj
6.8541 0 Td
(fish)Tj
1.6611 0 Td
(captured)Tj
3.7474 0 Td
(during)Tj
2.9253 0 Td
(the)Tj
1.4684 0 Td
(spring)Tj
-34.6278 -1.3039 Td
(2015)Tj
2.1203 0 Td
(were)Tj
2.1259 0 Td
(also)Tj
1.7632 0 Td
(recaptured)Tj
4.541 0 Td
(in)Tj
1.0375 0 Td
(the)Tj
1.4683 0 Td
(spring)Tj
2.7496 0 Td
(2016.)Tj
2.3471 0 Td
(All)Tj
1.3946 0 Td
(spring)Tj
2.7553 0 Td
(males)Tj
2.5001 0 Td
(either)Tj
2.5342 0 Td
(expelled)Tj
3.4923 0 Td
(sperm)Tj
2.7155 0 Td
(during)Tj
-33.5449 -1.2983 Td
(capture)Tj
3.2144 0 Td
(or)Tj
1.0999 0 Td
(had)Tj
1.7064 0 Td
(sperm)Tj
2.7156 0 Td
(removed)Tj
3.7474 0 Td
(via)Tj
1.3719 0 Td
(catheter)Tj
3.4242 0 Td
([)Tj
0.83 0.64 0.02 0 k
(18)Tj
0 g
(].)Tj
ET
Q
q
1 j
1 J
0 w
11.9999 0 0 11.9999 200.0125 425.0267 cm
BT
/F0 1 Tf
1 TL
-0.0033 Tc
0 0 Td
(Genetic)Tj
3.6283 0 Td
(analysis)Tj
ET
Q
q
1 j
1 J
0 w
10 0 0 10 200.0125 408.0188 cm
BT
/F6 1 Tf
1 TL
-0.004 Tc
0 0 Td
(Isolation)Tj
3.7303 0 Td
(methods,)Tj
3.9061 0 Td
(microsatellite)Tj
5.6069 0 Td
(loci,)Tj
1.8879 0 Td
(PCR)Tj
2.0636 0 Td
(conditions,)Tj
4.7055 0 Td
(visualization/scoring,)Tj
8.742 0 Td
(and)Tj
1.7291 0 Td
(locus)Tj
2.2847 0 Td
(verifi-)Tj
-34.6561 -1.3039 Td
(cation)Tj
2.6985 0 Td
(methods)Tj
3.685 0 Td
(all)Tj
1.1452 0 Td
(followed)Tj
3.651 0 Td
(those)Tj
2.3358 0 Td
(from)Tj
2.2053 0 Td
(Farrae)Tj
2.7836 0 Td
(et)Tj
0.9411 0 Td
(al.)Tj
1.1225 0 Td
([)Tj
0.83 0.64 0.02 0 k
(19)Tj
0 g
(].)Tj
-19.3718 -1.2982 Td
(COLONY)Tj
4.2859 0 Td
(2.0.6.2)Tj
2.8006 0 Td
(was)Tj
1.6951 0 Td
(used)Tj
2.0636 0 Td
(to)Tj
1.0205 0 Td
(identify)Tj
3.2995 0 Td
(full)Tj
1.5251 0 Td
(siblings)Tj
3.2428 0 Td
(prior)Tj
2.2563 0 Td
(to)Tj
1.0262 0 Td
(analysis)Tj
3.3221 0 Td
(with)Tj
/F11 1 Tf
2.0013 0 Td
[()10()7()-9(\r)4()-4()7(\r)12()20()]TJ
/F6 1 Tf
3.8097 0 Td
(because)Tj
-33.5449 -1.2983 Td
(having)Tj
2.9139 0 Td
(many)Tj
2.4775 0 Td
(closely)Tj
2.8857 0 Td
(related)Tj
2.9423 0 Td
(individuals)Tj
4.6488 0 Td
(can)Tj
1.6271 0 Td
(result)Tj
2.4491 0 Td
(in)Tj
1.0318 0 Td
(an)Tj
1.2075 0 Td
(overestimation)Tj
6.1965 0 Td
(of)Tj
1.0148 0 Td
(the)Tj
1.474 0 Td
(number)Tj
3.4129 0 Td
(of)Tj
1.0148 0 Td
(clus-)Tj
-35.2967 -1.3039 Td
(ters.)Tj
1.8992 0 Td
(All)Tj
1.3946 0 Td
(fish)Tj
1.6667 0 Td
(were)Tj
2.1204 0 Td
(analyzed)Tj
3.702 0 Td
(together)Tj
3.5376 0 Td
(first)Tj
1.8028 0 Td
(with)Tj
1.9956 0 Td
(three)Tj
2.2677 0 Td
(runs)Tj
2.0296 0 Td
(using)Tj
2.3867 0 Td
(the)Tj
1.4684 0 Td
(full)Tj
1.5307 0 Td
(likelihood)Tj
4.2349 0 Td
(and)Tj
1.7291 0 Td
(pairwise-)Tj
-33.766 -1.2983 Td
(likelihood)Tj
4.2349 0 Td
(combined)Tj
4.2463 0 Td
(method)Tj
3.3221 0 Td
(\(FPLS\))Tj
2.9991 0 Td
(with)Tj
1.9956 0 Td
(a)Tj
0.6576 0 Td
(polygamous)Tj
5.057 0 Td
(breeding)Tj
3.753 0 Td
(system,)Tj
3.1635 0 Td
(weak)Tj
2.262 0 Td
(priors,)Tj
2.8403 0 Td
(updat-)Tj
-34.5314 -1.2982 Td
(ing)Tj
1.5023 0 Td
(allele)Tj
2.2394 0 Td
(frequencies,)Tj
5.0059 0 Td
(and)Tj
1.7291 0 Td
(no)Tj
1.2756 0 Td
(genotyping)Tj
4.7168 0 Td
(errors)Tj
2.6249 0 Td
(assumed.)Tj
3.9005 0 Td
(One)Tj
1.9388 0 Td
(run)Tj
1.6668 0 Td
(was)Tj
1.7008 0 Td
(of)Tj
1.0148 0 Td
(medium)Tj
3.6056 0 Td
(length)Tj
2.7326 0 Td
(with)Tj
-35.6539 -1.2983 Td
(medium)Tj
3.6056 0 Td
(precision,)Tj
4.1329 0 Td
(one)Tj
1.6951 0 Td
(was)Tj
1.7008 0 Td
(of)Tj
1.0148 0 Td
(medium)Tj
3.6056 0 Td
(length)Tj
2.7269 0 Td
(with)Tj
2.0013 0 Td
(high)Tj
2.0069 0 Td
(precision,)Tj
4.1329 0 Td
(and)Tj
1.7291 0 Td
(one)Tj
1.6951 0 Td
(was)Tj
1.7008 0 Td
(of)Tj
1.0148 0 Td
(long)Tj
1.9899 0 Td
(length)Tj
-34.7525 -1.3039 Td
(with)Tj
1.9955 0 Td
(high)Tj
2.0069 0 Td
(precision.)Tj
4.1329 0 Td
(Each)Tj
2.177 0 Td
(spawning)Tj
4.0422 0 Td
(group)Tj
2.6079 0 Td
(was)Tj
1.7007 0 Td
(then)Tj
2.0183 0 Td
(analyzed)Tj
3.702 0 Td
(independently)Tj
5.9754 0 Td
(with)Tj
1.9956 0 Td
(a)Tj
0.6576 0 Td
(long)Tj
1.9843 0 Td
(run)Tj
-34.9963 -1.2983 Td
(length)Tj
2.7269 0 Td
(and)Tj
1.7291 0 Td
(high)Tj
2.0126 0 Td
(precision)Tj
3.9061 0 Td
(with)Tj
1.9956 0 Td
(all)Tj
1.1452 0 Td
(other)Tj
2.347 0 Td
(parameters)Tj
4.6772 0 Td
(the)Tj
1.4683 0 Td
(same.)Tj
2.4831 0 Td
(The)Tj
1.7802 0 Td
(independent)Tj
5.2781 0 Td
(COLONY)Tj
-31.5494 -1.2983 Td
(runs)Tj
2.0295 0 Td
(by)Tj
1.1792 0 Td
(spawning)Tj
4.0479 0 Td
(season)Tj
2.8516 0 Td
(were)Tj
2.126 0 Td
(also)Tj
1.7631 0 Td
(used)Tj
2.0637 0 Td
(to)Tj
1.0261 0 Td
(estimate)Tj
3.5433 0 Td
(effective)Tj
3.5092 0 Td
(population)Tj
4.5921 0 Td
(size)Tj
1.6838 0 Td
(\()Tj
/F11 1 Tf
[()]TJ
ET
Q
q
1 j
1 J
0 w
7.4999 0 0 6.6659 514.5448 276.3212 cm
BT
/F11 1 Tf
1 TL
-0.0053 Tc
0 0 Td
[()]TJ
ET
Q
q
1 j
1 J
0 w
10 0 0 10 517.4362 278.022 cm
BT
/F6 1 Tf
1 TL
-0.004 Tc
0 0 Td
(\))Tj
0.5612 0 Td
(for)Tj
1.389 0 Td
(a)Tj
0.6519 0 Td
(single)Tj
-34.3442 -1.3039 Td
(time)Tj
2.0296 0 Td
(point)Tj
2.3584 0 Td
(based)Tj
2.4718 0 Td
(on)Tj
1.2756 0 Td
(sibling)Tj
2.88 0 Td
(relationships)Tj
5.3517 0 Td
(among)Tj
2.9934 0 Td
(individuals.)Tj
-18.1643 -1.2982 Td
(The)Tj
1.7858 0 Td
(Bayesian)Tj
3.719 0 Td
(clustering)Tj
4.1556 0 Td
(program)Tj
/F11 1 Tf
3.7077 0 Td
[()10()7()-9(\r)4()-4()7(\r)12()20()]TJ
/F6 1 Tf
3.8097 0 Td
(2.3.4)Tj
2.0977 0 Td
([)Tj
0.83 0.64 0.02 0 k
(23)Tj
0 g
(])Tj
1.8538 0 Td
(was)Tj
1.6951 0 Td
(used)Tj
2.0636 0 Td
(to)Tj
1.0262 0 Td
(observe)Tj
3.2655 0 Td
(individual)Tj
4.2858 0 Td
(genetic)Tj
-34.6617 -1.2983 Td
(clustering)Tj
4.1499 0 Td
(over)Tj
1.9786 0 Td
(a)Tj
0.6519 0 Td
(range)Tj
2.4661 0 Td
(of)Tj
1.0148 0 Td
(populations)Tj
4.9493 0 Td
(\()Tj
/F11 1 Tf
[()]TJ
/F6 1 Tf
1.2019 0 Td
(varied)Tj
2.6985 0 Td
(15\).)Tj
2.2564 0 Td
(Run)Tj
1.9162 0 Td
(parameters)Tj
4.6715 0 Td
(were)Tj
2.126 0 Td
(set)Tj
1.3039 0 Td
(at)Tj
0.9523 0 Td
(10,000)Tj
2.8233 0 Td
(burn-)Tj
-35.1606 -1.3039 Td
(in)Tj
1.0318 0 Td
(repetitions)Tj
4.5071 0 Td
(followed)Tj
3.6453 0 Td
(by)Tj
1.1849 0 Td
(10,000)Tj
2.8232 0 Td
(Markov)Tj
3.3733 0 Td
(chain)Tj
2.4207 0 Td
(Monte-Carlo)Tj
5.4652 0 Td
(repetitions,)Tj
4.7281 0 Td
(with)Tj
1.9956 0 Td
(and)Tj
1.7347 0 Td
(without)Tj
-32.9099 -1.2983 Td
(location)Tj
3.4526 0 Td
(information)Tj
5.0739 0 Td
(included)Tj
3.7134 0 Td
(as)Tj
1.0148 0 Td
(a)Tj
0.652 0 Td
(prior,)Tj
2.4831 0 Td
(and)Tj
1.7291 0 Td
(with)Tj
1.9956 0 Td
(five)Tj
1.6554 0 Td
(independent)Tj
5.2724 0 Td
(runs)Tj
2.0296 0 Td
(per)Tj
/F11 1 Tf
1.5307 0 Td
[()]TJ
/F6 1 Tf
(.)Tj
1.0885 0 Td
(Each)Tj
2.1769 0 Td
(season)Tj
-33.868 -1.2982 Td
(also)Tj
1.7688 0 Td
(was)Tj
1.6951 0 Td
(evaluated)Tj
3.9912 0 Td
(independently)Tj
5.9754 0 Td
(under)Tj
2.6192 0 Td
(the)Tj
1.4683 0 Td
(same)Tj
2.262 0 Td
(conditions)Tj
4.4731 0 Td
(to)Tj
1.0261 0 Td
(identify)Tj
3.2938 0 Td
(any)Tj
1.6555 0 Td
(underlying)Tj
4.6033 0 Td
(struc-)Tj
-34.8318 -1.304 Td
(ture.)Tj
2.0636 0 Td
(Structure)Tj
3.9401 0 Td
(Harvester)Tj
4.1216 0 Td
([)Tj
0.83 0.64 0.02 0 k
(24)Tj
0 g
(])Tj
1.8538 0 Td
(was)Tj
1.7008 0 Td
(used)Tj
2.0579 0 Td
(to)Tj
1.0262 0 Td
(assess)Tj
2.5228 0 Td
(the)Tj
1.4683 0 Td
(results)Tj
2.8063 0 Td
(and)Tj
1.7348 0 Td
(determine)Tj
4.32 0 Td
(the)Tj
1.4683 0 Td
(number)Tj
3.4185 0 Td
(of)Tj
-34.503 -1.2982 Td
(groups)Tj
2.9707 0 Td
(that)Tj
1.7745 0 Td
(best)Tj
1.8084 0 Td
(fit)Tj
1.0772 0 Td
(the)Tj
1.4683 0 Td
(data)Tj
1.9163 0 Td
([)Tj
0.83 0.64 0.02 0 k
(25)Tj
0 g
(].)Tj
-9.8192 -1.2983 Td
(Genetic)Tj
3.3222 0 Td
(population)Tj
4.5921 0 Td
(structure)Tj
3.8211 0 Td
(within)Tj
2.8119 0 Td
(the)Tj
1.4683 0 Td
(James)Tj
2.5909 0 Td
(River)Tj
2.3584 0 Td
(\(spring-spawning)Tj
7.268 0 Td
(versus)Tj
2.7155 0 Td
(fall-spawning)Tj
-32.1446 -1.2983 Td
(sturgeon\))Tj
4.0592 0 Td
(was)Tj
1.7008 0 Td
(evaluated)Tj
3.9911 0 Td
(using)Tj
2.3868 0 Td
(pairwise)Tj
/F11 1 Tf
3.5489 0 Td
[()]TJ
ET
Q
q
1 j
1 J
0 w
7.4999 0 0 6.6659 362.0409 146.3244 cm
BT
/F11 1 Tf
1 TL
-0.0053 Tc
0 0 Td
[()20()]TJ
ET
Q
q
1 j
1 J
0 w
10 0 0 10 369.8645 148.0251 cm
BT
/F6 1 Tf
1 TL
-0.004 Tc
0 0 Td
(,)Tj
0.4422 0 Td
(a)Tj
0.6576 0 Td
(measure)Tj
3.5773 0 Td
(of)Tj
1.0148 0 Td
(genetic)Tj
3.0727 0 Td
(distance,)Tj
3.7304 0 Td
(as)Tj
1.0148 0 Td
(calculated)Tj
4.2009 0 Td
(in)Tj
-34.6957 -1.3039 Td
(GenAlEx)Tj
3.8721 0 Td
(6.502)Tj
2.347 0 Td
([)Tj
0.83 0.64 0.02 0 k
(26)Tj
0 g
(,)Tj
0.83 0.64 0.02 0 k
1.7348 0 Td
(27)Tj
0 g
(].)Tj
/F11 1 Tf
1.7405 0 Td
[()4()]TJ
ET
Q
q
1 j
1 J
0 w
7.4999 0 0 6.6659 304.1574 133.3417 cm
BT
/F11 1 Tf
1 TL
-0.0053 Tc
0 0 Td
[()20()]TJ
ET
Q
q
1 j
1 J
0 w
10 0 0 10 311.9811 134.9858 cm
BT
/F6 1 Tf
1 TL
-0.004 Tc
0 0 Td
(,)Tj
0.4422 0 Td
(a)Tj
0.6576 0 Td
(measure)Tj
3.5773 0 Td
(of)Tj
1.0148 0 Td
(genetic)Tj
3.0727 0 Td
(distance)Tj
3.5036 0 Td
(that)Tj
1.7802 0 Td
(is)Tj
0.8447 0 Td
(adjusted)Tj
3.5773 0 Td
(to)Tj
1.0261 0 Td
(the)Tj
1.4684 0 Td
(maximum)Tj
-32.1616 -1.2982 Td
(value)Tj
2.3017 0 Td
(of)Tj
/F11 1 Tf
1.0091 0 Td
[()]TJ
ET
Q
q
1 j
1 J
0 w
7.4999 0 0 6.6659 238.337 120.359 cm
BT
/F11 1 Tf
1 TL
-0.0053 Tc
0 0 Td
[()20()]TJ
ET
Q
q
1 j
1 J
0 w
10 0 0 10 248.3149 122.0031 cm
BT
/F6 1 Tf
1 TL
-0.004 Tc
0 0 Td
(for)Tj
1.3832 0 Td
(a)Tj
0.6463 0 Td
(given)Tj
2.3755 0 Td
(suite)Tj
2.0976 0 Td
(of)Tj
1.0091 0 Td
(markers)Tj
3.4923 0 Td
([)Tj
0.83 0.64 0.02 0 k
(28)Tj
0 g
(,)Tj
0.83 0.64 0.02 0 k
1.7348 0 Td
(29)Tj
0 g
(],)Tj
1.7348 0 Td
(was)Tj
1.6951 0 Td
(computed)Tj
4.2576 0 Td
(for)Tj
1.3833 0 Td
(the)Tj
1.4683 0 Td
(same)Tj
2.2564 0 Td
(pairwise)Tj
3.5376 0 Td
(compari-)Tj
-33.9021 -1.2982 Td
(son)Tj
1.6328 0 Td
(using)Tj
2.3924 0 Td
(GenAlEx)Tj
3.872 0 Td
(with)Tj
1.9956 0 Td
(999)Tj
1.6497 0 Td
(standard)Tj
3.7191 0 Td
(permutations)Tj
5.5955 0 Td
(and)Tj
1.7292 0 Td
(individual)Tj
4.2916 0 Td
(analysis)Tj
3.3222 0 Td
(suppressed.)Tj
-29.0038 -1.3039 Td
(Molecular)Tj
4.2972 0 Td
(diversity)Tj
3.6453 0 Td
(indices)Tj
3.0387 0 Td
(for)Tj
1.3833 0 Td
(the)Tj
1.4683 0 Td
(James)Tj
2.5909 0 Td
(River)Tj
2.3584 0 Td
(were)Tj
2.1203 0 Td
(calculated)Tj
4.2066 0 Td
(using)Tj
2.3867 0 Td
(GenAlEx,)Tj
4.0989 0 Td
(including)Tj
-32.7909 -1.2983 Td
(number)Tj
3.4186 0 Td
(of)Tj
1.0148 0 Td
(alleles)Tj
2.5964 0 Td
(per)Tj
1.5364 0 Td
(locus)Tj
2.2847 0 Td
(\()Tj
/F11 1 Tf
[()]TJ
ET
Q
q
1 j
1 J
0 w
7.4999 0 0 6.6659 318.8409 81.3543 cm
BT
/F11 1 Tf
1 TL
-0.0053 Tc
0 0 Td
[()]TJ
ET
Q
q
1 j
1 J
0 w
10 0 0 10 322.4692 82.9984 cm
BT
/F6 1 Tf
1 TL
-0.004 Tc
0 0 Td
(\),)Tj
0.788 0 Td
(allelic)Tj
2.5058 0 Td
(size)Tj
1.6838 0 Td
(range,)Tj
2.6872 0 Td
(and)Tj
1.7348 0 Td
(heterozygosity)Tj
5.9924 0 Td
(\()Tj
/F11 1 Tf
[()]TJ
ET
Q
q
1 j
1 J
0 w
7.4999 0 0 6.6659 487.1055 81.3543 cm
BT
/F11 1 Tf
1 TL
-0.0053 Tc
0 0 Td
[()]TJ
ET
Q
q
1 j
1 J
0 w
10 0 0 10 491.244 82.9984 cm
BT
/F6 1 Tf
1 TL
-0.004 Tc
0 0 Td
(\))Tj
0.5612 0 Td
([)Tj
0.83 0.64 0.02 0 k
(30)Tj
0 g
(].)Tj
2.075 0 Td
(Rarified)Tj
3.4185 0 Td
(allelic)Tj
ET
Q
q
1 j
1 J
0 w
576 737.1 m
36 737.1 l
36 737.6 l
576 737.6 l
f*
36 741.2598 107.1496 23.6976 re
W* n
q
107.0929 0 0 23.6409 36 741.3165 cm
q
/I0 Do
Q
Q
Q
q
0 0 612 792 re
W* n
1 j
1 J
0 w
7.9999 0 0 7.9999 317.3669 745.7952 cm
BT
/F0 1 Tf
1 TL
-0.005 Tc
0 0 Td
(Genetic)Tj
3.6071 0 Td
(differentiation)Tj
6.1087 0 Td
(of)Tj
1.0417 0 Td
(male)Tj
2.3457 0 Td
(Atlantic)Tj
3.4441 0 Td
(sturgeon)Tj
4.0394 0 Td
(in)Tj
0.978 0 Td
(the)Tj
1.5874 0 Td
(James)Tj
3.1252 0 Td
(River,)Tj
2.7851 0 Td
(Virginia)Tj
ET
Q
q
1 j
1 J
0 w
36 48.0002 m
576 48.0002 l
576 47.5002 l
36 47.5002 l
f*
0.83 0.64 0.02 0 k
81.5811 34.9228 m
237.8835 34.9228 l
h
f*
0 g
7.9999 0 0 7.9999 36 36 cm
BT
/F0 1 Tf
1 TL
-0.005 Tc
0 0 Td
(PLOS)Tj
2.8559 0 Td
(ONE)Tj
2.3669 0 Td
(|)Tj
0.83 0.64 0.02 0 k
0.4748 0 Td
(https://doi.or)Tj
5.4142 0 Td
(g/10.137)Tj
3.8198 0 Td
(1/journal.po)Tj
5.1236 0 Td
(ne.01796)Tj
4.089 0 Td
(61)Tj
0 g
2.0906 0 Td
(July)Tj
1.9701 0 Td
(7,)Tj
1.0417 0 Td
(2017)Tj
36.4397 0 Td
(3)Tj
0.7654 0 Td
(/)Tj
ET
endstream
endobj
98 0 obj
<>stream
endstream
endobj
99 0 obj
<>stream
endstream
endobj
100 0 obj
<>stream
endstream
endobj
101 0 obj
<>stream
endstream
endobj
102 0 obj
<>stream
endstream
endobj
103 0 obj
<>stream
endstream
endobj
104 0 obj
<>stream
endstream
endobj
105 0 obj
<>stream
endstream
endobj
106 0 obj
<>stream
endstream
endobj
107 0 obj
<>stream
endstream
endobj
108 0 obj
<>stream
endstream
endobj
109 0 obj
<>stream
endstream
endobj
110 0 obj
<>stream
BT
66.9477 0 Td
(8)Tj
ET
Q
endstream
endobj
111 0 obj
<>/XObject<>>>/CropBox[0 0 612 792]/MediaBox[0 0 612 792]/Parent 13 0 R/Annots 113 0 R/Contents 114 0 R/TrimBox[0 0 612 792]>>
endobj
112 0 obj
<>
endobj
115 0 obj
<>stream
/CIDInit/ProcSet findresource begin
12 dict begin begincmap
/CMapName/Adobe-Identity-UCS def
/CIDSystemInfo<>def
/CMapType 2 def
1 begincodespacerange
<0000>
endcodespacerange
1 beginbfchar
<00EF><0394>
endbfchar
endcmap
CMapName currentdict /CMap defineresource pop end end
endstream
endobj
116 0 obj
<>/FontDescriptor 117 0 R/DW 250/W[239[603]]>>
endobj
117 0 obj
<>
endobj
118 0 obj
<>stream
cvt f6 fpgm2Msf bglyfcW Dhead= 6hhea1 $hmtx&O